Refereed Journal Articles
1. Proctor, H. and G. Pritchard. 1989. Neglected predators: water mites (Acari: Parasitengona: Hydrachnellae) in freshwater communities. Journal of the North American Benthological Society 8: 100-111.
2. Proctor, H.C. 1989. Occurrence of protandry and a female-biased sex-ratio in a sponge-associated water mite (Acari: Unionicolidae). Experimental and Applied Acarology 7: 289-297.
3. Proctor, H.C. and G. Pritchard. 1990. Variability in the life history of Unionicola crassipes, a sponge-associated water mite. Canadian Journal of Zoology 68: 1227-1232.
4. Proctor, H.C. and G. Pritchard. 1990. Prey detection by the water mite Unionicola crassipes (Acari: Unionicolidae). Freshwater Biology 23: 271-290.
5. Proctor, H.C. 1991. Courtship in the water mite Neumania papillator: males capitalize on female adaptations for predation. Animal Behaviour 42:589-598.
6. Proctor, H.C. 1991. The evolution of copulation in water mites: a comparative test for nonreversing characters. Evolution 45: 558-567.
7. Johnson, D.L. and H.C. Proctor. 1991. Effects of phytoseiid predators on the sex ratio of the spider mite Panonychus ulmi. Canadian Journal of Zoology 69: 208-212.
8. Proctor, H.C. 1992. Mating and spermatophore morphology of water mites (Acari: Parasitengona). Zoological Journal of the Linnean Society 106: 341-384.
9. Proctor, H.C. 1992. Discord between field and laboratory sex ratios of the water mite Neumania papillator Marshall (Acari: Unionicolidae). Canadian Journal of Zoology 70: 2483-2486.
10. Proctor, H.C. 1992. Effect of food deprivation on mate-searching and spermatophore production in male water mites (Acari: Unionicolidae). Functional Ecology 6: 661-665.
11. Proctor, H.C. 1992. Cohabitation of six species of spiders in webs of Cyrtophora moluccensis (Araneae, Araneidae) in Moorea, French Polynesia. Journal of Arachnology 20: 144-145.
12. Proctor, H.C. 1992. Sensory exploitation and the evolution of male mating behaviour: a cladistic test using water mites (Acari: Parasitengona). Animal Behaviour 44: 745-752.
13. Baker, R.L., M.R.L. Forbes and H.C. Proctor. 1992. Sexual differences in development and behaviour of larval Ischnura verticalis (Odonata: Coenagrionidae). Canadian Journal of Zoology 70: 1161-1165.
14. Proctor, H.C. 1993. Mating biology resolves trichotomy for cheliferoid pseudoscorpions (Pseudoscorpionida, Cheliferoidea). The Journal of Arachnology 21: 156-158.
15. Proctor, H.C. and L.D. Harder. 1994. Pollen load, capsule weight and seed production in three orchid species. Canadian Journal of Botany. 72: 249-255.
16. Proctor, H.C. and B.P. Smith. 1994. Mating behaviour of the water mite Arrenurus manubriator (Acari: Arrenuridae). Journal of Zoology, London 232: 473-483.
17. Proctor, H.C., R.L. Baker, and D.T. Gwynne. 1995. Mating behaviour and spermatophore morphology: a comparative test of the female choice hypothesis. Canadian Journal of Zoology, 73: 2010-2020.
18. Proctor, H.C. and L.D. Harder. 1995. Effect of pollination success on floral longevity in the orchid Calypso bulbosa (Orchidaceae). American Journal of Botany, 82: 1131-1136.
19. Proctor, H.C. 1997. Mating behavior of Physolimnesia australis (Halik) (Acari: Limnesiidae), a non-parasitic, rotifer-eating water mite from Australia. Journal of Arachnology 25: 321-325.
20. Proctor, H.C., H.M. Gray and B.M. OConnor. 1997. Subaquatic mites (Acari: Astigmata) associated with adult freshwater leeches (Hirudinea: Erpobdellidae). Journal of Natural History 31: 539-544.
21. Garga, N., H. Proctor and R. Belczewski. 1997. Leg size affects mating success in Tarsonemus confusus Ewing (Prostigmata: Tarsonemidae). Acarologia 38: 369-375.
22. Walter, D.E. and H. C. Proctor. 1998. Feeding behaviour and phylogeny: observations on early derivative Acari. Experimental and Applied Acarology 22: 39-50.
23. Proctor, H.C. 1998. Indirect sperm transfer in arthropods. Annual Review of Entomology 43: 153-174.
24. Proctor, H.C. 1998. Effect of pollen age on fruit set, fruit weight, and seed set in three orchid species. Canadian Journal of Botany 76: 420-427.
25. Walter, D.E. and H.C. Proctor. 1998. Predatory mites in tropical Australia: local species richness and complementarity. Biotropica 30: 72-81.
26. Proctor, H.C. 1999. Gallilichus jonesi sp. n. (Acari: Ascouracaridae), a new species of feather mite from the quills of the Australian Brush-Turkey (Aves: Megapodiidae). Australian Journal of Entomology, 38: 77-84.
27. Proctor, H.C. and I. Owens. 2000. Mites and birds: diversity, parasitism and coevolution. Trends in Ecology and Evolution 15: 358-364.
28. Proctor, H.C. 2001. Extracting aquatic mites from stream substrates: a comparison of three methods. Experimental and Applied Acarology 25: 1-11.
29. Proctor, H.C. 2001. Megninia casuaricola sp. n. (Acari: Analgidae), the first feather mite from a cassowary (Aves: Struthioniformes: Casuariidae). Australian Journal of Entomology 40: 335-341.
30. Williams, C.R. and H.C. Proctor. 2002. Parasitism of mosquitoes (Diptera: Culicidae) by larval mites (Acari: Parasitengona) in Adelaide, South Australia. Australian Journal of Entomology 41: 161-163.
31. Göth, A and H. Proctor. 2002. Pecking preferences in hatchlings of the Australian brush-turkey, Alectura lathami (Megapodiidae): the role of food type and colour. Australian Journal of Zoology 50: 93-102.
32. Proctor, H., K. Montgomery, K. Rosen and R.L. Kitching. 2002. Are tree trunks habitats or highways? A comparison of oribatid mite assemblages from hoop-pine bark and litter. Australian Journal of Entomology 41: 294-299.
33. Proctor, H.C. 2003. Feather mites (Acari: Astigmata): ecology, behavior and evolution. Annual Review of Entomology 48: 185-209.
34. Nakamura, A., H. Proctor and C.P. Catterall. 2003. Using soil and litter arthropods to assess the state of rainforest restoration. Ecological Management & Restoration 4 (supplement): S20-S28.
35. Mironov, S.V., J. Dabert and H. Proctor. 2003. New feather mites of the family Pterolichidae (Acari: Pterolichoidea) from parrots (Aves: Psittaciformes) in Australia. Australian Journal of Entomology 42: 185-202.
36. Fan Q.-H., Walter D.E. and Proctor H.C. 2003. A review of the family Barbutiidae (Acari: Raphignathoidea), with the description of two new species from Australia. Systematic & Applied Acarology 8: 107-130.
37. Kanowski, J., Catterall, C. P., Wardell-Johnson, G. W., Proctor, H. and Reis, T. 2003. Development of forest structure on cleared rainforest land in eastern Australia under different styles of reforestation. Forest Ecology and Management 183: 265-280.
38. Fan, Q.-H., Walter, D.E., and Proctor, H. 2003. A review of the genus Ledermuelleriopsis Willmann (Acari: Prostigmata: Stigmaeidae). Invertebrate Systematics 17: 551-574.
39. Proctor, H.C., J. Kanowski, G. Wardell-Johnson, T. Reis and C.P. Catterall. 2003. Does diversity beget diversity? A comparison between plant and leaf-litter invertebrate richness from pasture to rainforest. Records of the South Australian Museum Monograph Series No. 7: 267-274.
40. Proctor, H.C. and Jones, D.N. 2004. Geographical structuring of feather mite assemblages from the Australian brush-turkey (Aves: Megapodiidae). Journal of Parasitology 90: 60-66.
41. Boulton, A., M. Harvey and H. Proctor. 2004. Of spates and species: responses by interstitial water mites to simulated spates in a subtropical Australian river. Experimental and Applied Acarology 34: 149-169.
42. Proctor, H.C. and N. Garga. 2004. Red, distasteful water mites: did fish make them that way? Experimental and Applied Acarology 34: 127-147.
43. Lian Pin Koh, R.R. Dunn, N.S. Sodhi, R.K. Colwell, H.C. Proctor, and V.S. Smith. 2004. Species coextinctions and the biodiversity crisis. Science 305: 1632-1634.
44. Mironov, S.V. and H.C. Proctor. 2005. A new feather mite genus of the family Psoroptoididae (Acari: Analgoidea) from cassowaries. Journal of Natural History 39: 3297-3304.
45. Bochkov, A.V., G. Williams and H. Proctor. 2005. First record of Picobia zumpti (Acari: Syringophilidae) from quills of the rock pigeon in North America and description of the male. Belgian Journal of Entomology 7: 121-127.
46. Proctor, H and A. Grigg. 2006. Aquatic invertebrates in final void water bodies at an open-cut coal mine in central Queensland. Australian Journal of Entomology 45: 107-121.
47. Dabert, J., S.V. Mironov and H. Proctor. 2006. A new species of the genus Titanolichus Gaud & Atyeo, 1996 (Acari: Astigmata: Pterolichidae) from the endangered orange-bellied parrot Neophema chrysogaster (Aves: Psittaciformes) from Australia. Australian Journal of Entomology 45: 206-214.
48. Beaulieu, F., D. E. Walter, H. C. Proctor, R. L. Kitching and F. Menzel. 2006. Mesostigmatid mites (Acari: Mesostigmata) on rainforest tree trunks: arboreal specialists, but substrate generalists? Experimental and Applied Acarology 39: 25-40.
49. Knee, W. and H. Proctor. 2006. Keys to the families and genera of blood and tissue feeding mites associated with Albertan birds. Canadian Journal of Arthropod Identification 2: 1-18.
50. Proctor, H., G. Zimmerman and K. Meyer. 2006. A new feather mite, Aetacarus elanoides sp. n. (Acari: Gabuciniidae), from the Swallow-tailed Kite Elanoides forficatus (Linnaeus) (Falconiformes: Accipitridae; Perninae). Zootaxa 1252: 37-47.
51. Grimbacher, P.S., C.P. Catterall, J. Kanowski and H.C. Proctor. 2007. Responses of ground-active beetle assemblages to different styles of reforestation on cleared rainforest land. Biodiversity and Conservation 16: 2167-2184.
52. Boeckner, M. and H. Proctor. 2005*. Water-bears from the Rocky Mountains: a first look at Alberta's tardigrade fauna. Canadian Field Naturalist 119: 586-588 [*note: although issue is dated 2005, it was not actually published until May 2007]
53. Knee, W. and H. Proctor. 2007. Host records for the northern fowl mite, Ornithonyssus sylviarum (Mesostigmata: Macronyssidae), from birds of North America (Canada, USA, and Mexico). Journal of Medical Entomology 44: 709-713.
54. Mironov, S.V., H.C. Proctor, M. Barreto and G. Zimmerman. 2007. New genera and species of feather mites of the family Gabuciniidae (Astigmata: Pterolichoidea) from New World raptors (Aves: Falconiformes). The Canadian Entomologist 139: 757-777.
55. Kelly, J.R., H. Proctor and J.P. Volpe. 2008. Intertidal community structure differs significantly between substrates dominated by native eelgrass (Zostera marina L.) and adjacent to the introduced oyster Crassostrea gigas (Thunberg) in British Columbia, Canada. Hydrobiologia 596: 57-66.
56. Gentes, M.-L., H. Proctor and G. Wobeser. 2007. Demodicosis in a mule deer (Odocoileus hemionus hemionus) from Saskatchewan, Canada. Journal of Wildlife Diseases 43: 758–761.
57. Galván, I., E. Barba, R. Piculo, J. L. Cantó, V. Cortés, J. S. Monrós, F. Atiénzar and H. Proctor. 2008. Feather mites and birds: an interaction mediated by uropygial gland size? Journal of Evolutionary Biology 21: 133-144.
58. Knee, W., H. Proctor and T. Galloway. 2008. Survey of nasal mites (Rhinonyssidae, Ereynetidae, and Turbinoptidae) associated with birds in Alberta and Manitoba, Canada. The Canadian Entomologist 140: 364-379.
59. Mironov, S.V. and H.C. Proctor. 2008. The probable association of feather mites of the genus Ingrassia (Analgoidea: Xolalgidae) with the blue penguin Eudyptula minor (Aves: Spenisciformes) in Australia. Journal of Parasitology 94: 1243-1248.
61. Williams, B.W., S.R. Gelder and H. Proctor. 2009. Distribution and first reports of Branchiobdellida (Annelida: Clitellata) on crayfish in the Prairie Provinces of Canada. Western North American Field Naturalist 69: 119-124.
62. Boeckner, M.J., J. Sharma and H.C. Proctor. 2009. Revisiting the meiofauna paradox: dispersal and colonization of nematodes and other meiofaunal organisms in low- and high-energy environments. Hydrobiologia: 624: 91–106.
63. Proctor, H. 2009. Can freshwater mites act as forensic tools? Experimental and Applied Acarology 49: 161-165.
64. Mironov, S. and H. Proctor. 2009. Feather mites of the genus Proterothrix Gaud (Astigmata: Proctophyllodidae) from parrotbills (Passeriformes: Paradoxornithidae) in China. Journal of Parasitology: 95: 1093-1107.
65. Stephenson, S., S. Hannon and H. Proctor. 2009. The function of feathers in tree swallow nests: insulation or ectoparasite barrier? The Condor 111: 479-487.
66. Mironov, S. and H. Proctor. 2009. Atelophyllodes gen. n., a new feather mite genus of the family Proctophyllodidae (Astigmata: Analgoidea) from lyrebirds (Passeriformes: Menuridae). Zootaxa 2326: 51–61.
67. Knee, W. and H. Proctor. 2010. Interactive HTML-based dichotomous key to female Rhinonyssidae (Mesostigmata) from birds in Canada. Canadian Journal of Arthropod Identification 9 (doi: 10.3752/cjai.2010.09: available online at http://www.biology.ualberta.ca/bsc/ejournal/kp09/kp_09.html ).
68. Walter, D.E. and H.C. Proctor. 2010. Mites as modern models: acarology in the 21st century. Acarologia 50: 131-141 (invited paper for Acarologia’s 50th anniversary).
69. Beaulieu, F., D.E. Walter, H.C. Proctor and R.L. Kitching. 2010. The canopy starts at 0.5 m: predatory mites (Acari: Mesostigmata) differ between rain forest floor soil and suspended soil at any height. Biotropica 42: 704-709.
70. Kontschán, J., H. Proctor and J. Newton. 2010. Trachyuropoda kinsella n. sp. (Acari: Uropodina: Trachyuropodidae) from Alberta, Canada, with a key to Trachyuropoda species from temperate North America. International Journal of Acarology 36: 211-220.
71. Carleton, R.E. and H.C. Proctor. 2010. Feather mites associated with Eastern Bluebirds (Sialia sialis L.) in Georgia, including the description of a new species of Trouessartia (Analgoidea: Trouessartiidae). Southeastern Naturalist 9: 605-623.
72. Haribal, M. , H. Proctor, A.A. Dhondt, and E. Rodriguez. 2011. Biology of House Finch feather mites, Proctophyllodes pinnatus (Acari: Proctophyllodidae), parallels variation in preen gland secretions. International Journal of Acarology 37: 75 – 90.
73. Mori, B.A., H.C. Proctor, D.E. Walter and M.L. Evenden. 2011. Phoretic mite associates of mountain pine beetle at the leading edge of an infestation in northwestern Alberta, Canada. The Canadian Entomologist 143: 44-55.
74. Mestre, A., F. Mesquita-Joanes, H. Proctor and J. S. Monrós. 2011. Different scales of spatial segregation of two species of feather mites on the wings of a passerine bird. Journal of Parasitology 97: 237-244.
75. Gale, K.S.P. and H.C. Proctor. 2011. Diets of two congeneric species of crayfish worm (Annelida: Clitellata; Branchiobdellidae) from western Canada. Canadian Journal of Zoology 89: 289-296.
76. Williams, B.W., H.C. Proctor and T. Clayton. 2011. Range extension of the Northern Crayfish, Orconectes virilis (Decapoda, Cambaridae), in the western prairie provinces of Canada. Crustaceana 84: 451-460.
77. Valim, M.P., F.A. Hernandes and H.C. Proctor. 2011. Feather mites of Brazil (Acari: Astigmata: Analgoidea and Pterolichoidea). International Journal of Acarology 37: 293-324.
78. Williams, B.W., K.L. Williams, S.R. Gelder and H.C. Proctor. 2011. Distribution of Entocytheridae (Crustacea: Ostracoda) in the northern prairies of North America and reports of opportunistic clitellate annelids on crayfish hosts. Western North American Naturalist 71: 276-282.
79. Mironov, S.V. and H.C. Proctor. 2011. Four new feather mite species of the family Pteronyssidae (Astigmata: Analgoidea) from laughing-thrushes (Passeriformes: Timaliidae) in China. Acarina 19: 35-51.
80. Newton, J.S., J. Glasier, H.E.L. Maw, H.C. Proctor and R.G. Foottit. 2011. Ants and subterranean Sternorrhyncha in a native grassland in east-central Alberta, Canada. The Canadian Entomologist 143: 518-523.
81. Locke, S.A, A.R. Lapierre, K. Byers, H. Proctor, J.D. McLaughlin and D.J. Marcogliese. 2012. Molecular and morphological evidence for the Holarctic distribution of Urogonimus macrostomus (Rudolphi, 1803) Monticelli, 1888 (Digenea: Leucochloridiidae). Journal of Parasitology 98: 880-882.