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A. Richard Palmer, FRSC
Professor
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Dept. of Biological Sciences
University of Alberta
Edmonton, Alberta T6G 2E9
    CANADA
 
Z1116 Biological Sciences Bldg
+1 (780) 492-3633, 3308
+1 (780) 492-9234
rich.palmer@ualberta.ca

| Research / Teaching | People & Projects | Publications | CV |
| Many views of Biological Asymmetry |
| Computer Programs | Data files | Links |
| New CSZ Section: Comparative Morphology & Development |
"The shadow of the bamboo sweeps the stair all night long,
yet not a mote of dust is stirred.
The moonbeams penetrate to the bottom of the pool,
yet in the water not a trace is left."

Chikan Zenji, circa AD 850

Current Research
  Evolution by natural selection requires three steps. New phenotypic variation 1) must arise, 2) must have an impact on fitness (survival or fecundity), and 3) must (ultimately) be heritable. The first step - how new variation arises - remains controversial. Traditionally, new phenotypes are ascribed to novel genotypes (mutants or recombinants). But developmental plasticity - the same genotype yields different forms in different environments - may be a much more important source of new phenotypes than generally recognized. This has renewed interest in its evolutionary significance.

Our work has yielded valuable insights into the causes and adaptive significance of several striking examples of developmental plasticity and has advanced some of the strongest evidence to date for a 'phenotype-precedes-genotype' mode of evolution. We continue to explore the interplay between developmental plasticity and evolution on both ecological time scales (via descriptive and experimental studies) and evolutionary time scales (via comparative studies).

  Ongoing research includes:
  • developmental plasticity in starfish tube feet.
  • handedness and induced asymmetry in crab claws.
  • learning and induced morphological defense in intertidal snails.
  • developmental plasticity in feeding limbs of barnacles and porcelain crabs.
  • development and evolution of conspicuous biological asymmetries in many taxa, including experimental studies of larval caddisflies and young Cancer crabs, and further comparative studies of these and other taxa (see People & Projects).

Most of our research is done at the Bamfield Marine Sciences Centre, a magnificent field station on the west coast of Vancouver Island.

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Our research is funded by:

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Current Teaching
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B361.GIF, 10K

 
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ZOOL250
Survey of Invertebrates

(extensive WWW content)
BIOL361
Marine Science
BIOL506
Systematics & Evolution Forum
MASC415
Animal Structure & Function

(in the Bamfield Fall Program)
Professional Honors

  • Invited Participant US National Academy of Sciences Colloquium: Symmetries Throughout the Sciences (June 1996)

  • McCalla Research Professor
    University of Alberta (2000/2001)

  • Faculty of Science Award for Excellent Teaching, University of Alberta (2000/2001)

  • Invited Plenary Lecture Gordon Research Conference: Organic Reactions & Processes (July 2006)
76th President (1995)
Western Society of Naturalists
Presidential address: "On koans and two-headed fish"
Invited Plenary Lecture (2000)
German Zoological Society Annual Meeting, Bonn, Germany
Invited Plenary Lecture (2001)
International Symposium on Chirality, Orlando, Florida
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Invited Plenary Lecture (2003)
100th Anniversary University of Puerto Rico Graduate Student Research Symposium: "Biology of the New Millennium", San Juan, Puerto Rico

Inaugural Chair (2005-2007)
Comparative Morphology & Development Section of the Canadian Society of Zoologists

Fellow (Elected 2007),
Royal Society of Canada
(UA Express News Story)
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Graduate Students and Post-Doctoral Fellows
Publications and Computer Programs
  SciCov.jpg, 14K Download pdf's of selected publications or view abstracts and figures.   Claw Landmarks.GIF, 3K Macintosh computer programs for morphometric data collection.   MacCoil.GIF, 2K Macintosh computer programs for teaching.  
Links

 

Systematics and Evolution Home Page.
Biological Sciences Home Page
Copyright (c) 1997 - 2008 by A. Richard Palmer. All rights reserved.
(revised Oct. 24, 2008)

have come before (since Jan. 1, 1997)