Clade for PLATYHELMINTHES
Clade for classes and Turbellarian orders from Nielson'94 p. 216.
CL. TUR # ==========2a*==B=============# Or. Acoela <<=1*==| E | | =======2c*===L===========# Or. Polycladida =2b*| L | | ====2e*====A=========# Or. Tricladida (planarians) =2d*| R | | ==2g*====IA======# Or. Rhabocoela =2f*| | ======4*==== Cl. Trematoda (endoparasitic flukes) ==3*==| | ==6*== Monogenea (ectoparasites of aquatic vertebrates) ==5*==| ==7*== Cl. Cestoidea (endoparasitic tapeworms)
1: a) midventral mouth**, b) blind gut**, c) protonephridia**, d) coelom & blood vascular system absent**,
e) ciliated epidermis**, f) multiciliated cells, g) epidermal gland cells extensive, h) monociliated sperm,
i) hermaphroditic (unclear if valid trait), j) archoopheran reproductive system,
2a: a) lack gut**, b) simple pharynx**, c) lack protonephridia, d) no distinct gonads (gametes from parenchyma)
2b: a) rhabdites, b) biciliated sperm (present in all higher groups), c) gut cavity present, d) plicate pharynx,
2c: a) gut has multiple branches, b) pair of anterior tentacles (some), c) multiple gonads among branches of gut
2d: a) neoopheran reproductive system (present in all higher groups); b) lose typical 'spiral' cleavage
2e: a) gut has three branches
2f: a) bulbous pharynx,
2g: a) gut has single cavity
3: a) syncytial, non-ciliated epidermis, b) terminal mouth (also some rhabdocoels), c) paired intestinal cecae
4: a) oral & mid-ventral sucker, b) complex=indirect life cycle (1-2 intermediate hosts), c) anaerobic metabolism
5: a) distinct attachment hooks in larva
6: a) haptor, b) simple=direct life cycle** (no intermediate hosts)
7: a) scolex, b) strobila (=chain of proglottids), c) mouth & gut absent, d) reproductive system repeated in each proglottid, e) complex=indirect life cycle (1 or more intermediate hosts),
** possibly plesiomorphic (a primitive state, not unique to clade)
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Copyright © 1997 by A. Richard Palmer. All rights reserved. (revised Feb. 3, 1997)